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Please help me understand Iolokus plot

Okay...I feel like an idiot even asking this. I understand Rousch is all about the human experimentation and whatnot, but I'm having trouble understanding the whole thing behind Mulder's brothers, who Jason is (besides being Mulder's brother, of course), what the hell happened to Samantha, etc. I need someone to really dumb it down for me. I'm almost done reading all four books for the very first time, and I'd like to read them all again in a couple of months...I'm sure I'll catch more the second time around, but any explanation would be greatly appreciated.

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